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They have a limited pressure release function when closing the doors and when moving will likely have limited cabin ventilation function by sitting in a negative pressure area, both despite the two fairly air impervious membranes both in the vent insert and roof lining itself.
I've never known water ingress through them; that panel does have propensity to rot, but mainly stemming from the friction fitting clips holding the vents in place and the rear lip trim in place that scratch the paint...
If your panel is rotten, the club do a GRP replacement panel which is supplied without the vent apertures having been cut out...
That was pretty much the cause of all the rot on my boot and roof. My solution, after repairs, was to pack the trim with grease to keep the water out.
Way back in the late-1970s or early-1980s, when I noticed rust in the areas around the push-on spring-clips (which scraped off the underlying paint) for the trim of my 1974 Triumph Toledo, I removed both before refurbishing the underlying steel & paintwork, but never replaced the trim owing to its propensity for retaining water and dirt. Those areas have never since rusted!
Although it would nice to reinstate the trim, I would be disinclined to do so, unless a reasonable way could be found that would not encourage corrosion again!?!
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Regards.
Nigel A. Skeet
Independent tutor of mathematics, physics, technology & engineering, for secondary, tertiary, further & higher education.
https://www.linkedin.com/profile/view?id=308177758
Upgraded 1974 Triumph Toledo 1300 (Toledo / Dolomite HL / Sprint hybrid)
Onetime member + magazine editor & technical editor of Volkswagen Type 2 Owners' Club